Yungas paramo grasslands (Puina, 2015).

The fourth study site, in the yungueño paramo grasslands between 3,600 and 4,100 m a.s.l., is a humid and cold region dominated by large grass species (in some cases more than 1.5 m high), with the ground covered by mosses, lichens and small ferns. There are also isolated keñua trees that are the only ones that grow at this altitude, forming small stands known as bosquesillos de keñua, which are threatened at a continental scale.

In total, 34 plant species, 9 species and subspecies of diurnal butterflies and 75 vertebrate species were recorded, of which 18 are new species for Madidi, 1 for Bolivia and 4 for science.